There are lots of degraded oak savannas, especially
in the hill country of southwestern Wisconsin. Most of these
savannas were grazed but were never plowed or logged. Often
many of the original open-grown bur oak trees are still
present, usually on the ridge tops, but sometimes in the
draws or on the south-facing slopes. If the savanna was
fairly far from the barn, it may not even have been heavily
grazed.
If there are no open-grown oaks left, restoration
is impossible. This is because it takes 100-150 years to
get the oaks back. But if open-grown oaks are present, the
savanna can be restored! In fact, it may only take a dozen
years or so to bring back the savanna understory.
Here are some principles for restoring a degraded
savanna:
-
- Remove invasive trees (black walnut, elm, etc.) that are
crowding the open-grown oaks. This is what is called "daylighting
the oaks." Treat
all cut stumps with herbicide to prevent resprouting.
If there are young oak trees, leave them.
- Remove all invasive brush, such as honeysuckle, buckthorn,
prickly ash, gray dogwood, brambles, etc., by cutting. Treat
all cut stumps with herbicide. Native shrubs such as
hazel and viburnum can be left. They will be top-killed
by fire but will resprout. However, leave only scattered
native shrubs; a dense shrub layer is undesirable.
- Burn. If there are good
oaks present there should be enough fuel on the ground,
in the form of oak leaves, to carry a fire.
- Monitor in the first year after the burn to see what comes
up. You may be surprised! At Pleasant Valley we find lots
of shooting star popping up (see photo below). They were
there all along but did not flower because they were too
shaded.

At this point, two options are available.
1) Annual burning followed by handplanting with a good mix
of savanna understory species. This is the approach we have
used at Pleasant Valley Conservancy (there had been no grazing
in at least 50 years). Burning followed by seeding must be
repeated at least for several years. Since fall planting is
usually best, burn in the fall. Burning will eliminate oak
leaves and open up the ground so that planted seeds can reach
the mineral soil. Plant savanna specialist species, as many
kinds as possible. Table of Brian Pruka's
list of savanna indicator species. Some species can be
planted throughout, but in especially sunny or shady areas
plant species better adapted to those habitats. Plant savanna
grasses in patches or swaths rather than uniformly throughout.
If only small amounts of seeds of certain species are available,
plant those in favored locations.
2) If the understory has been recently grazed, there will
probably be no desirable understory left. If this is the case,
use a heavy herbicide treatment to destroy whatever groundlayer
is present and start from scratch. This is the approach that
Don Schmidt used in a 5-acre savanna in the Town of Dunn,
Dane County, Wisconsin. This
is the approach that prairie restorationists use on agricultural
land. Depending on topgraphy and other conditions, herbicide
treatment might be possible with a tractor-operated boom sprayer.
If this is not possible, use a sprayer hose or hand-held boom
sprayer connected to an herbicide tank in a truck. An electric
or gasoline-powered pump at the tank provides pressure. However,
if heavy equipment is used, avoid driving over the roots of
the large trees. Make sure the whole understory is sprayed.
After spraying, either burn (if there is enough fuel) or rake
the area to expose the bare ground before seeding. Use
the same savanna species list as in option #1.
Because of the heavy work involved, clearing the savanna
is usually a multiple-year job. It should "not"
be done by conventional loggers, as they have no appreciation
for the delicacy of the woods. There are commercial restoration
companies that specialize in restoring savannas. However,
even if such a company does the work, the local site manager
must monitor the work carefully to be certain that desirable
trees are not cut. Unless the manager has considerable experience
with and confidence in the contractor, each tree to be cut
should be clearly marked. Brush and trees should be removed
at the same time. In Wisconsin, the time of year for this
work is winter, preferably when there is snow on the ground.
With snow present, fire can be used to eliminate brush, small
trees, and branches of large trees.
Whether option #1 or #2 is followed will depend on the size
of the restoration, the extent of degradation, the funds available,
etc. In either case, the critical item is the availability
of a good mix of savanna understory species. Don't start
a restoration until seeds are available! You need to get the
understory started right after clearing, before invasive shrubs
or brambles from the seed bank start to fluorish.
Some species may appear the first summer after planting,
but others may take four or five years.
It is important to monitor the area being restored frequently,
three or four times in a growing season. Mowing or spot spraying
may be necessary to knock back undesirable species.
Brian
Pruka's List of Savanna Indicator Species
The table below gives a list
of savanna indicator species that was developed by Brian Pruka.
This is Pruka's Category #1 list, species which are the best
indicators of former savannas and open woodlands. "These
are species that tend to be limited to partial canopy conditions.
In more densely-wooded sites, these species are usually in
a state of declining vigor due to the ever-increasing canopy
closure. They typically persist in densely-wooded sites only
near canopy openings such as woodlot edges, roadsides, or
brushed utility corridors." This list was published in
the 1995 Midwest Oak Savanna and Woodland Ecosystems Conference,
sponsored by the Environmental Protection Agency. Pruka's
paper should be consulted for further detail, and for additional
lists. A more detailed list which also includes extensive
background on savanna species can be found in Bader, Brian
J. 2001. Ecological Restoration 19:
242-250.
Not all these species will be
available, but the higher diversity the better. Note that
since this list was published, some of the Latin names have
been changed, although the common names mostly remain the
same.
| Latin name |
Common name |
| Actaea pachypoda |
White baneberry |
| Agastache nepetoides |
Yellow giant hyssop |
| Agastache scrophularieaefolia |
Purple giant hyssop |
| Agrimonia gryposepala |
Tall agrimony |
| Agropyron trachycaulum |
Slender wheatgrass |
| Agrostis perennans |
Autumn bentgrass |
| Allium cernuum |
Nodding wild onion |
| Amorpha canescens |
Lead-plant |
| Amphicarpa bracteata |
Hog peanut |
| Andropogon gerardi |
Big bluestem |
| Andropogon scoparius |
Little bluestem |
| Anemone cylindrica |
Thimbleweed |
| Anemone patens |
Pasque flower |
| Anemone quinquefolia |
Wood anemone |
| Anemone virginiana |
Tall anemone |
| Anemonellla thalictroides |
Rue anemone |
| Apocynum androsaemifolium |
Spreading dogbane |
| Apocynum cannabinum |
Indian hemp |
| Arenaria lateriflora |
Wood sandwort |
| Asclepias amplexicaulis |
Sand milkweed |
| Asclepias exaltata |
Poke milkweed |
| Asclepias purpurascens |
Purple milkweed |
| Asclepias tuberosa |
Butterfly milkweed |
| Asclepias viridiflora |
Short green milkweed |
| Asplenium platyneuron |
Ebony spleenwort |
| Aster azureus |
Sky-blue aster |
| Aster laevis |
Smooth blue aster |
| Aster linariifolias |
Flax-leaved aster |
| Aster sericeus |
Silky aster |
| Aster shortii |
Short's aster |
| Astragalus canadensis |
Canadian milkvetch |
| Aureolaria grandiflora |
Yellow false foxglove |
| Aureolaria pedicularia |
Clammy false foxglove |
| Baptisia leucantha |
White wild indigo |
| Baptisia leucophaea |
Cream wild indigo |
| Besseya bullii |
Kitten tails |
| Blephilia ciliata |
Ohio horse-mint |
| Bouteloua curtipendula |
Side oats grama |
| Bouteloua hirsuta |
Hairy grama grass |
| Brachyelytrum erectum |
Long-awned wood grass |
| Bromus ciliatus |
Fringed brome |
| Bromus kalmii |
Prairie brome |
| Bromus purgans |
Woodland brome |
| Cacalia atriplicifolia |
Pale Indian plantain |
| Cacalia muhlenbergii |
Great Indian plantain |
| Cacalia suaveolens |
Sweet Indian plantain |
| Callirhoe triangulata |
Clustered poppy mallow |
| Camassia scilloides |
Wild hyacinth |
| Carex brevior |
Sedge |
| Carex rugosperma |
a sedge |
| Cassia fasciculata |
Partridge pea |
| Castilleja occinea |
Indian paintbrush |
| Ceanothus americana |
New Jersey tea |
| Ceanothus ovatus |
Prairie redroot |
| Cirsium altissimum |
Woodland thistle |
| Comandra umbellata |
False toadflax |
| Convolvulus spithamaeus |
Low bindweed |
| Corallorhiza odontorhiza |
Late coral-root |
| Coreopsis palmata |
Prairie tickseed |
| Cypripedium pubescens |
Large yellow lady-slipper |
| Desmodium canadense |
Showy tick-trefoil |
| Desmodium canescens |
a tick-trefoil |
| Desmodium cuspidatum |
Bracted tick-trefoil |
| Dodecatheon meadia |
Shooting star |
| Elymus villosus |
Silky wild rye |
| Elymus virginicus |
Virginia wild rye |
| Erigeron pulchellus |
Robin's plantain |
| Eupatorium purpureum |
Purple joe-pye weed |
| Eupatorium sessilifolium |
Woodland boneset |
| Euphorbia corollata |
Flowering spurge |
| Galium boreale |
Northern bedstraw |
| Galium circaezans |
Wild licorice |
| Galium concinnum |
Shining bedstraw |
| Gentiana alba |
Cream gentian |
| Gentiana puberulenta |
Downy gentian |
| Gentiana quinquefolia |
Stiff gentian |
| Geum triflorum |
Prairie smoke |
| Helianthemum bicknellii |
Rockrose |
| Helianthemum canadense |
Common rockrose |
| Helianthus decapetalus |
Pale sunflower |
| Helianthus hirsutus |
Oblong sunflower |
| Helianthus laetiflorus |
Prairie sunflower |
| Helianthus occidentalis |
Western sunflower |
| Heuchera americana |
Prairie alum-root |
| Hieracium candense |
Canada hawkweed |
| Hieracium longipilum |
Hairy hawkweed |
| Hieracium scabrum |
Rough hawkweed |
| Hypoxis hirsuta |
Yellow star-grass |
| Hystrix patula |
Bottlebrush grass |
| Koeleria cristata |
June grass |
| Krigia biflora |
False dandelion |
| Kuhnia eupatorioides |
False boneset |
| Lactuca floridana |
Woodland blue lettuce |
| Lactuca ludoviciana |
Western prairie lettuce |
| Lathyrus ochroleucus |
Pale pea |
| Lathyrus venosus |
Woodland pea |
| Lechea intermedia |
Intermediate pinweed |
| Lechea tenuifolia |
Slender-leaved pinweed |
| Leersia virginica |
White grass |
| Lespedeza violacea |
Violet bush clover |
| Liatris aspera |
Rough blazing star |
| Liatris cylindracea |
Cylindrical blazing star |
| Liatris ligulistylis |
Showy blazing star |
| Liatris pychnostachya |
Prairie blazing star |
| Lilium philadelphicum |
Prairie lily |
| Lithospermum canescens |
Hoary puccoon |
| Lithospermum caroliniense |
Hairy puccoon |
| Lithospermum incisum |
Fringed puccoon |
| Lobelia spicata |
Pale spiked lobelia |
| Luzula multiflora |
Common wood-rush |
| Lysimachia lanceolata |
Lance-leaved loosestrife |
| Lysimachia quadriflora |
Narrow-leaved loosestrife |
| Lysimachia quadrifolia |
Whorled loosestrife |
| Oenothera perennis |
Small sundrops |
| Orobanche uniflora |
One-flowered broom-rape |
| Oryzopis racemosa |
Black-seeded ricegrass |
| Oxalis violacea |
Violet wood-sorrel |
| Panicum lanuginosum var. implicatum |
Hairy panic-grass |
| Panicum latifolium |
Broad-leaved panic-grass |
| Panicum linearifolium |
Slender-leaved panic-grass |
| Panicum oligosanthes var. scribnerianum |
Few-flowered panic-grass |
| Pedicularis canadensis |
Wood betony |
| Penstemon gracilis |
Slender beard-tongue |
| Penstemon pallidus |
Pale beard-tongue |
| Petalostemum candidum |
White prairie clover |
| Petalostemum purpureum |
Purple prairie clover |
| Phlox pilosa |
Prairie phlox |
| Polemonium reptans |
Jacob's ladder |
| Polygala sanguinea |
Field milkwort |
| Polygala senega |
Seneca snakeroot |
| Polygala verticillata |
Whorled milkwort |
| Polytaenia nutallii |
Prairie parsley |
| Potentilla arguta |
Prairie cinquefoil |
| Prenanthes alba |
Lion's foot |
| Pycnanthemum virginianum |
Common mountain mint |
| Ranunculus fascicularis |
Early buttercup |
| Ranunculus rhomboideus |
Prairie buttercup |
| Ratibida pinnata |
Yellow coneflower |
| Rudbeckia hirta |
Black-eyed Susan |
| Silene antirrhina |
Sleepy catchfly |
| Silene stellata |
Starry campion |
| Sisyrinchium campestre |
Prairie blue-eyed grass |
| Smilacina stellata |
Starry false Solomon's seal |
| Solidago hispida |
Hairy goldenrod |
| Solidago juncea |
Early goldenrod |
| Solidago nemoralis |
Old-field goldenrod |
| Solidago rigida |
Stiff goldenrod |
| Solidago speciosa |
Showy goldenrod |
| Solidago ulmifolia |
Elm-leaved golden rod |
| Sorghastrum nutans |
Indian grass |
| Sporobolus heterolepis |
Prairie dropseed grass |
| Symphoricarpos occidentalis |
Wolfberry |
| Taenidia integerrima |
Yellow pimpernel |
| Tephrosia virginiana |
Goat's rue |
| Thaspium trifoliatum |
Meadow parsnip |
| Tomanthera auriculata |
Eared false foxglove |
| Tradescantia ohiensis |
Common spiderwort |
| Trillium recurvatum |
Red trillium |
| Triosteum aurantiacum |
Late horse gentian |
| Triosteum perfoliatum |
Early horse gentian |
| Uvularia grandiflora |
Bellwort |
| Veronicastrum virginicum |
Culver's root |
| Vicia americana |
American vetch |
| Vicia caroliniana |
Wood vetch |
| Viola pedata |
Bird's foot violet |
| Viola pedatifida |
Prairie violet |
| Zigadenus elegans |
White camas |
| Zizia aurea |
Golden Alexander |
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