Oak Savanna Restoration

Bringing Back a Degraded Oak Savanna

There are lots of degraded oak savannas, especially in the hill country of southwestern Wisconsin. Most of these savannas were grazed but were never plowed or logged. Often many of the original open-grown bur oak trees are still present, usually on the ridge tops, but sometimes in the draws or on the south-facing slopes. If the savanna was fairly far from the barn, it may not even have been heavily grazed.

If there are no open-grown oaks left, restoration is impossible. This is because it takes 100-150 years to get the oaks back. But if open-grown oaks are present, the savanna can be restored! In fact, it may only take a dozen years or so to bring back the savanna understory.

Here are some principles for restoring a degraded savanna:

  • Determine that open-grown oaks are present. (Look for the typical open-grown character.)
  • Remove invasive trees (black walnut, elm, etc.) that are crowding the open-grown oaks. This is what is called "daylighting the oaks." Treat all cut stumps with herbicide to prevent resprouting. If there are young oak trees, leave them.
  • Remove all invasive brush, such as honeysuckle, buckthorn, prickly ash, gray dogwood, brambles, etc., by cutting. Treat all cut stumps with herbicide. Native shrubs such as hazel and viburnum can be left. They will be top-killed by fire but will resprout. However, leave only scattered native shrubs; a dense shrub layer is undesirable.
  • Burn. If there are good oaks present there should be enough fuel on the ground, in the form of oak leaves, to carry a fire.
  • Monitor in the first year after the burn to see what comes up. You may be surprised! At Pleasant Valley we find lots of shooting star popping up (see photo below). They were there all along but did not flower because they were too shaded.

 

At this point, two options are available.

1) Annual burning followed by handplanting with a good mix of savanna understory species. This is the approach we have used at Pleasant Valley Conservancy (there had been no grazing in at least 50 years). Burning followed by seeding must be repeated at least for several years. Since fall planting is usually best, burn in the fall. Burning will eliminate oak leaves and open up the ground so that planted seeds can reach the mineral soil. Plant savanna specialist species, as many kinds as possible. Table of Brian Pruka's list of savanna indicator species. Some species can be planted throughout, but in especially sunny or shady areas plant species better adapted to those habitats. Plant savanna grasses in patches or swaths rather than uniformly throughout. If only small amounts of seeds of certain species are available, plant those in favored locations.

2) If the understory has been recently grazed, there will probably be no desirable understory left. If this is the case, use a heavy herbicide treatment to destroy whatever groundlayer is present and start from scratch. This is the approach that Don Schmidt used in a 5-acre savanna in the Town of Dunn, Dane County, Wisconsin. This is the approach that prairie restorationists use on agricultural land. Depending on topgraphy and other conditions, herbicide treatment might be possible with a tractor-operated boom sprayer. If this is not possible, use a sprayer hose or hand-held boom sprayer connected to an herbicide tank in a truck. An electric or gasoline-powered pump at the tank provides pressure. However, if heavy equipment is used, avoid driving over the roots of the large trees. Make sure the whole understory is sprayed. After spraying, either burn (if there is enough fuel) or rake the area to expose the bare ground before seeding. Use the same savanna species list as in option #1.

Because of the heavy work involved, clearing the savanna is usually a multiple-year job. It should "not" be done by conventional loggers, as they have no appreciation for the delicacy of the woods. There are commercial restoration companies that specialize in restoring savannas. However, even if such a company does the work, the local site manager must monitor the work carefully to be certain that desirable trees are not cut. Unless the manager has considerable experience with and confidence in the contractor, each tree to be cut should be clearly marked. Brush and trees should be removed at the same time. In Wisconsin, the time of year for this work is winter, preferably when there is snow on the ground. With snow present, fire can be used to eliminate brush, small trees, and branches of large trees.

Whether option #1 or #2 is followed will depend on the size of the restoration, the extent of degradation, the funds available, etc. In either case, the critical item is the availability of a good mix of savanna understory species. Don't start a restoration until seeds are available! You need to get the understory started right after clearing, before invasive shrubs or brambles from the seed bank start to fluorish.

Some species may appear the first summer after planting, but others may take four or five years.

It is important to monitor the area being restored frequently, three or four times in a growing season. Mowing or spot spraying may be necessary to knock back undesirable species.


Brian Pruka's List of Savanna Indicator Species

The table below gives a list of savanna indicator species that was developed by Brian Pruka. This is Pruka's Category #1 list, species which are the best indicators of former savannas and open woodlands. "These are species that tend to be limited to partial canopy conditions. In more densely-wooded sites, these species are usually in a state of declining vigor due to the ever-increasing canopy closure. They typically persist in densely-wooded sites only near canopy openings such as woodlot edges, roadsides, or brushed utility corridors." This list was published in the 1995 Midwest Oak Savanna and Woodland Ecosystems Conference, sponsored by the Environmental Protection Agency. Pruka's paper should be consulted for further detail, and for additional lists. A more detailed list which also includes extensive background on savanna species can be found in Bader, Brian J. 2001. Ecological Restoration 19: 242-250.

Not all these species will be available, but the higher diversity the better. Note that since this list was published, some of the Latin names have been changed, although the common names mostly remain the same.

Latin name Common name
Actaea pachypoda White baneberry
Agastache nepetoides Yellow giant hyssop
Agastache scrophularieaefolia Purple giant hyssop
Agrimonia gryposepala Tall agrimony
Agropyron trachycaulum Slender wheatgrass
Agrostis perennans Autumn bentgrass
Allium cernuum Nodding wild onion
Amorpha canescens Lead-plant
Amphicarpa bracteata Hog peanut
Andropogon gerardi Big bluestem
Andropogon scoparius Little bluestem
Anemone cylindrica Thimbleweed
Anemone patens Pasque flower
Anemone quinquefolia Wood anemone
Anemone virginiana Tall anemone
Anemonellla thalictroides Rue anemone
Apocynum androsaemifolium Spreading dogbane
Apocynum cannabinum  Indian hemp
Arenaria lateriflora Wood sandwort
Asclepias amplexicaulis Sand milkweed
Asclepias exaltata Poke milkweed
Asclepias purpurascens Purple milkweed
Asclepias tuberosa Butterfly milkweed
Asclepias viridiflora Short green milkweed
Asplenium platyneuron Ebony spleenwort
Aster azureus Sky-blue aster
Aster laevis Smooth blue aster
Aster linariifolias Flax-leaved aster
Aster sericeus Silky aster
Aster shortii Short's aster
Astragalus canadensis Canadian milkvetch
Aureolaria grandiflora Yellow false foxglove
Aureolaria pedicularia Clammy false foxglove
Baptisia leucantha White wild indigo
Baptisia leucophaea Cream wild indigo
Besseya bullii Kitten tails
Blephilia ciliata Ohio horse-mint
Bouteloua curtipendula Side oats grama
Bouteloua hirsuta Hairy grama grass
Brachyelytrum erectum Long-awned wood grass
Bromus ciliatus Fringed brome
Bromus kalmii Prairie brome
Bromus purgans Woodland brome
Cacalia atriplicifolia Pale Indian plantain
Cacalia muhlenbergii Great Indian plantain
Cacalia suaveolens Sweet Indian plantain
Callirhoe triangulata Clustered poppy mallow
Camassia scilloides Wild hyacinth
Carex brevior Sedge
Carex rugosperma a sedge
Cassia fasciculata Partridge pea
Castilleja occinea Indian paintbrush
Ceanothus americana New Jersey tea
Ceanothus ovatus Prairie redroot
Cirsium altissimum Woodland thistle
Comandra umbellata False toadflax
Convolvulus spithamaeus Low bindweed
Corallorhiza odontorhiza Late coral-root
Coreopsis palmata Prairie tickseed
Cypripedium pubescens Large yellow lady-slipper
Desmodium canadense Showy tick-trefoil
Desmodium canescens a tick-trefoil
Desmodium cuspidatum Bracted tick-trefoil
Dodecatheon meadia Shooting star
Elymus villosus Silky wild rye
Elymus virginicus  Virginia wild rye
Erigeron pulchellus Robin's plantain
Eupatorium purpureum  Purple joe-pye weed
Eupatorium sessilifolium Woodland boneset
Euphorbia corollata Flowering spurge
Galium boreale Northern bedstraw
Galium circaezans Wild licorice
Galium concinnum Shining bedstraw
Gentiana alba Cream gentian
Gentiana puberulenta Downy gentian
Gentiana quinquefolia Stiff gentian
Geum triflorum  Prairie smoke
Helianthemum bicknellii Rockrose
Helianthemum canadense Common rockrose
Helianthus decapetalus Pale sunflower
Helianthus hirsutus Oblong sunflower
Helianthus laetiflorus Prairie sunflower
Helianthus occidentalis Western sunflower
Heuchera americana Prairie alum-root
Hieracium candense Canada hawkweed
Hieracium longipilum Hairy hawkweed
Hieracium scabrum Rough hawkweed
Hypoxis hirsuta Yellow star-grass
Hystrix patula Bottlebrush grass
Koeleria cristata June grass
Krigia biflora False dandelion
Kuhnia eupatorioides False boneset
Lactuca floridana Woodland blue lettuce
Lactuca ludoviciana Western prairie lettuce
Lathyrus ochroleucus Pale pea
Lathyrus venosus Woodland pea
Lechea intermedia  Intermediate pinweed
Lechea tenuifolia Slender-leaved pinweed
Leersia virginica White grass
Lespedeza violacea Violet bush clover
Liatris aspera  Rough blazing star
Liatris cylindracea Cylindrical blazing star
Liatris ligulistylis Showy blazing star
Liatris pychnostachya Prairie blazing star
Lilium philadelphicum Prairie lily
Lithospermum canescens Hoary puccoon
Lithospermum caroliniense Hairy puccoon
Lithospermum incisum Fringed puccoon
Lobelia spicata Pale spiked lobelia
Luzula multiflora Common wood-rush
Lysimachia lanceolata Lance-leaved loosestrife
Lysimachia quadriflora Narrow-leaved loosestrife
Lysimachia quadrifolia Whorled loosestrife
Oenothera perennis Small sundrops
Orobanche uniflora One-flowered broom-rape
Oryzopis racemosa Black-seeded ricegrass
Oxalis violacea Violet wood-sorrel
Panicum lanuginosum var. implicatum Hairy panic-grass
Panicum latifolium Broad-leaved panic-grass
Panicum linearifolium Slender-leaved panic-grass
Panicum oligosanthes var. scribnerianum Few-flowered panic-grass
Pedicularis canadensis Wood betony
Penstemon gracilis Slender beard-tongue
Penstemon pallidus Pale beard-tongue
Petalostemum candidum White prairie clover
Petalostemum purpureum Purple prairie clover
Phlox pilosa Prairie phlox
Polemonium reptans Jacob's ladder
Polygala sanguinea Field milkwort
Polygala senega Seneca snakeroot
Polygala verticillata Whorled milkwort
Polytaenia nutallii Prairie parsley
Potentilla arguta Prairie cinquefoil
Prenanthes alba Lion's foot
Pycnanthemum virginianum Common mountain mint
Ranunculus fascicularis Early buttercup
Ranunculus rhomboideus Prairie buttercup
Ratibida pinnata Yellow coneflower
Rudbeckia hirta Black-eyed Susan
Silene antirrhina Sleepy catchfly
Silene stellata Starry campion
Sisyrinchium campestre Prairie blue-eyed grass
Smilacina stellata Starry false Solomon's seal
Solidago hispida Hairy goldenrod
Solidago juncea Early goldenrod
Solidago nemoralis Old-field goldenrod
Solidago rigida Stiff goldenrod
Solidago speciosa Showy goldenrod
Solidago ulmifolia Elm-leaved golden rod
Sorghastrum nutans Indian grass
Sporobolus heterolepis Prairie dropseed grass
Symphoricarpos occidentalis Wolfberry
Taenidia integerrima Yellow pimpernel
Tephrosia virginiana Goat's rue
Thaspium trifoliatum Meadow parsnip
Tomanthera auriculata Eared false foxglove
Tradescantia ohiensis Common spiderwort
Trillium recurvatum Red trillium
Triosteum aurantiacum Late horse gentian
Triosteum perfoliatum Early horse gentian
Uvularia grandiflora Bellwort
Veronicastrum virginicum Culver's root
Vicia americana American vetch
Vicia caroliniana Wood vetch
Viola pedata Bird's foot violet
Viola pedatifida Prairie violet
Zigadenus elegans White camas
Zizia aurea Golden Alexander

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